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Genomic comparisons and the adaptive basis of brain size plasticity and chromosomal instability in the Eurasian common shrew

  • William R. Thomas
  • , Tanya M. Lama
  • , Cecilia Baldoni
  • , Laia Marín-Gual
  • , Diana Moreno Santillán
  • , Marta Farré
  • , Linelle Abueg
  • , Jennifer Balacco
  • , Olivier Fedrigo
  • , Giulio Formenti
  • , Nivesh Jain
  • , Jacquelyn Mountcastle
  • , Tatiana Tilley
  • , Ying Sims
  • , Alan Tracey
  • , Jo Wood
  • , David A. Ray
  • , Dominik Von Elverfeldt
  • , John Nieland
  • , Angelique P. Corthals
  • Aurora Ruiz-Herrera, Dina K.N. Dechmann, Erich Jarvis, Liliana M. Dávalos
  • Stony Brook University
  • Smith College
  • Max Planck Institute of Animal Behavior
  • University of Konstanz
  • Autonomous University of Barcelona
  • University of California at Berkeley
  • University of Kent
  • Rockefeller University
  • Wellcome Sanger Institute
  • Texas Tech University
  • University of Freiburg
  • Aalborg University
  • City University of New York
  • Howard Hughes Medical Institute

Research output: Contribution to journalArticlepeer-review

1 Scopus citations

Abstract

Sorex araneus, the Eurasian common shrew, has seasonal brain size plasticity (Dehnel's phenomenon) and many intraspecific chromosomal rearrangements. Genomic contributions to these traits, however, remain unknown. We couple a chromosome-scale genome assembly with seasonal brain transcriptomes to discover relationships between molecular evolution and both traits. While Positively Selected Genes (PSGs) enriched the Fanconi anemia DNA repair pathway (FANCI, FAAP100), which is likely involved in chromosomal rearrangements by preventing the accumulation of chromosomal aberrations, genes under positive selection or showing seasonal differential expression in the brain implicate neurogenesis (PCDHA6, SOX9, Notch signaling) and metabolic regulation (VEGFA, SPHK2) as key mechanisms underlying Dehnel's phenomenon. We also find that both positively selected and differentially expressed genes in the hippocampus are overrepresented near S. araneus evolutionary breakpoints. This relates both positive selection and differential expression to accessible chromatin configuration, suggesting that chromosomal rearrangements are integral to adaptive evolution and the regulation of brain size plasticity.

Original languageEnglish
Article numbermsag006
JournalMolecular Biology and Evolution
Volume43
Issue number2
DOIs
StatePublished - Feb 1 2026

Keywords

  • Dehnel’s phenomenon
  • chromosomal evolution
  • cortex
  • evolutionary breakpoints
  • highly contiguous genome assembly
  • hippocampus
  • shrew

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